A vida complexa possui células complexas, também conhecidas como células eucaria³ticas. Ao contra¡rio das bactanãrias, por exemplo, as células da vida complexa tem muitos compartimentos internos distintos conhecidos como organelas.
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)
Fig. 1 A reticulação em esferas BS 3 revela um complexo altamente interligado. (A) Purificação de afinidade em tandem de TPC usando subunidades marcadas com TML e AtEH1 / Pan1. O complexo purificado foi analisado por coloração com prata em SDS-PAGE de 4 a 20%. Todas as subunidades TPC, exceto LOLITA, foram identificadas com base no MS (arquivo de dados S8) e são indicadas entre os dois ganãis. M, marcador; HSP70, proteana de choque tanãrmico 70. EF1, fator de alongamento 1; TEV, Tobacco Etch Virus. (B) TML-GS purificado por afinidade em tandem antes e depois da reticulação com várias concentrações de BS 3, analisado por coloração com prata em um gel de SDS-PAGE de 4 a 20%. O vasto tamanho do TPC, com um peso molecular esperado de 914 kDa, émanifestado pela perda de subunidades individuais e o acaºmulo de proteanas incapazes de penetrar no gel de empilhamento. (C) Ana¡lise de reticulação após purificação em tandem de subunidades marcadas com TML e AtEH1 / Pan1 expressas em culturas de células PSB-D visualizadas por Xvis. Cada análise se origina de um total de seis experimentos e combina conjuntos de dados BS 3 de 1,2 e 5 mM . (D) Um exemplo do espectro de fragmentação da reticulação intersubunidade entre TWD40-1 e TWD40-2, conforme indicado em (C). Crédito: Science Advances (2021). DOI: 10.1126 / sciadv.abe7999
As células dependem de membranas para se proteger do mundo exterior. Mas as membranas celulares não podem ser totalmente fechadas porque os nutrientes e outras moléculas precisam ser capazes de passar. Para conseguir isso, as membranas celulares tem muitos tipos de canais e poros. Além disso, existem receptores embutidos na membrana que monitoram continuamente o mundo externo e sinalizam para o interior da canãlula. A colaboração extensiva entre cinco grupos VIB resultou em uma melhor compreensão da maquinaria que as plantas usam para regular a composição proteica de sua membrana externa. Esta descoberta, publicada na Science Advances, aumenta nosso conhecimento ba¡sico de como a composição da membrana plasma¡tica pode ser adaptada com base em estamulos externos, um processo essencial para a vida na Terra.
A arquitetura molecular do TPLATE
A vida complexa possui células complexas, também conhecidas como células eucaria³ticas. Ao contra¡rio das bactanãrias, por exemplo, as células da vida complexa tem muitos compartimentos internos distintos conhecidos como organelas. Essas organelas trocam material entre si. Para isso, as organelas possuem alguns truques, um deles éo tra¡fego de vesaculas. Isso significa que eles usam uma parte de sua própria membrana como uma bolsa para as mercadorias a serem trocadas.
Uma descoberta recente mostrou que as plantas dependem muito de um complexo de proteanas chamado complexo TPLATE para fazer isso. Este complexo não estãoapenas presente em plantas, mas também em uma ampla gama de outros eucariotos, o que sugere que éevolutivamente antigo e faz parte de uma familia de complexo de proteanas da qual todos os outros membros são intensamente estudados. No entanto, como esse complexo particular não estãopresente nos organismos modelo mais estudados (animais e leveduras), sua existaªncia e função permaneceram por muito tempo sob o radar.
Neste estudo, as equipes do VIB revelaram a arquitetura molecular do TPLATE pela primeira vez. Eles conseguiram isso reticulando espectrometria de massa e simulações de computador. Essas novas percepções revelaram a orientação desse complexo em direção a membrana , bem como a delicada relação entre os diferentes domanios de suas subunidades.
Esses achados são importantes para aumentar o conhecimento dos processos eucaria³ticos cruciais. Na verdade, a estrutura desse complexo agora nos permite compara¡-lo com estruturas conhecidas de seus parentes pra³ximos que estãopresentes em todos os eucariotos, incluindo animais e leveduras, permitindo aos pesquisadores visualizar a evolução desses complexos de tra¡fico.
Para obter uma visão estrutural e funcional deste complexo enigma¡tico, era necessa¡ria uma abordagem integrativa e colaborativa. Cinco grupos de pesquisa do VIB e um grupo da República Tcheca contribuaram com sua experiência especafica para realizar experimentos que variam de estudos de ligação de lipadios a abordagens de biologia estrutural.
O novo insight estrutural foi gerado principalmente com base em espectrometria de massa de reticulação , realizada com a ajuda da instalação de núcleo VIB Proteomics.
"Um grande benefacio de trabalhar no VIB éque ele incentiva e facilita muito o acesso ao conhecimento e experiência que permite que grupos de pesquisa embarquem com sucesso em projetos conjuntos que estãomuito além de sua zona de conforto", disse o Prof.
Este estudo servira¡ de base para futuros trabalhos cientaficos e abrira¡ portas para a geração de novos e mais seguros herbicidas ou para a modulação das respostas ao estresse nas plantas.